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The Motion Path feature makes it possible to precisely control the path that a particle takes after being emitted. This is done by setting keyframes for a comp Light and naming the Light in a certain way.
This keyframed Light lets you control the path that particles travel after being born. Motion Path is useful when precise control of the particle path is needed.
The popup for Motion Path is the only Particular parameter that you will set when creating a motion path. Each of the numbers corresponds a number 'X'. That number is used in the name given to the comp Light that acts as the motion source.